Wildlife Impacts on Agroecosystems and Culture: Achieving Integrated Pest Management of Invasive Ungulates in Hawai‘i

Progress report for SW22-935

Project Type: Research and Education
Funds awarded in 2022: $349,979.00
Projected End Date: 07/31/2025
Grant Recipient: University of Hawaii
Region: Western
State: Hawaii
Principal Investigator:
Dr. Melissa Price
University of Hawaii
Co-Investigators:
Kyle Caires
University of Hawaii
Derek Risch
University of Hawaii
Stephanie Shwiff
USDA National Wildlife Research Center, Dept of Economics
John Steensma
Steensma Dairy
Dr. Mark Thorne
University of Hawaii at Manoa
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Project Information

Summary:

Hawai'i lacks native ungulates, but introductions of hoofed mammals have resulted in population explosions of wild ungulates on each of the islands in the archipelago (Figure 1). Aside from domesticated livestock, these nonnative ungulates are feral, invasive, or both (Table 1). Although ungulates are a potentially unifying issue linking diverse stakeholders in meeting common challenges regarding food security and environmental health, the needs of ranchers, farmers, state wildlife managers, and hunters have largely been viewed as competing, rather than complementary. State agency management of game in Hawai‘i, in contrast to the continental states, does not extend to private lands, creating a tremendous management burden for producers on private lands. Meanwhile, producers on state lease lands find themselves in a position of accommodating hunting access and regulations that may directly hinder their operations or prevent them from optimizing conservation practices. By identifying individual landowners and managers as decision makers for land parcels, while simultaneously addressing issues of land adjacency and potential synergistic solutions amongst stakeholders, we are contributing to collaborative change in ungulate management to increase stability and sustainability for agriculture in Hawai‘i. In this project we determined relationships between ecological and economic health of ranch and agricultural lands, identified densities of unmanaged ungulates, and developed data-driven Integrated Pest Management (IPM) decision guidelines to improve on-farm wildlife management and profitability.

Related WSARE projects have successfully quantified economic damages from wildlife to livestock (Elser et al 2019) and highlighted benefits of native wildlife to agroecosystems (Smith and El-Swaify 2006; Price et al 2021). Negative impacts of unmanaged ungulates are well-documented in native, non-agricultural ecosystems (Leopold & Hess 2017), but prior to this project, economic damage and forage loss from wild nonnative ungulates inhabiting agricultural lands had not been quantified in the Hawaiian Islands.

In this project we surveyed for ungulate presence and impacts using economic, social, and ecological criteria. The economic component of this project demonstrated that wild unmanaged ungulates are major pests in the Hawaiian Islands, decreasing productivity and seriously threatening the viability of ranching in Hawai‘i. Further, the field component of this research demonstrated that more than 60% of sites had a decrease in forage due to invasive unmanaged ungulates, and a number of sites had a substantial amount of bare ground due to ongoing drought conditions compounded by overgrazing by wild ungulates. While not quantified in this study, we received reports of a number of small ranchers ceasing ranching activities or even selling their ranches due to these combined impacts of drought and wild ungulates, and most ranchers reported reductions in herd size.

We note that wild ungulates are still valued by both hunters and producers for recreation, cultural practices, and food security (Conover 1998, Lohr et al. 2014, Luat-Hūʻeu et al. 2021, Luat-Hūʻeu et al. 2023). Thus, in this project we also identified management actions that improve ranch production, profitability and accessibility, reduce pests, and increase native biodiversity, while respecting cultural values. To achieve these integrated values, our project leadership team includes native Hawaiians, hunters, ranchers, conservationists. Further, our research is co-developed with native Hawaiian practitioners and experts, and we work alongside agricultural and natural resource management experts to implement, interpret, and publish our research. Thus, the experimental design, the implementation of research, and the interpretation of results are guided by native Hawaiians who are experts in ranching and land management.

Over the last three years, a combination of commercial harvest, fencing, targeted hunts by community members and land managers, and aerial shooting by land managers has substantially curbed the exponential growth that Axis deer were previously experiencing. A considerable increase in investments in deer fencing by the state legislature, counties, USDA, and other grants has reduced the movement of Axis deer (and other wild ungulates) between regions, decreasing impacts in some places but also increasing impacts to some places where removal efforts have not yet been intensified. Ranches are working with community hunters where feasible to reduce Axis deer and Mouflon sheep impacts to ranches while feeding communities. Commercial hunting, while largely limited to a single operator, has been hugely effective in knocking down populations while demonstrating a viable business model. Funding provided by the state to support aerial shooting knockdown efforts, where requested by ranchers, has been implemented by state agencies, who hold the only permits for such activities. Thus, despite impacts from devastating wildfires and ongoing droughts during the project period, substantial progress has been made in increased collaboration among land managers, funding for fencing infrastructure, and a deeper understanding regarding the level of effort needed to minimize impacts to ranches from invasive unmanaged ungulates.

These substantial reductions in Axis deer abundance are expected to improve forage, crop, and soil quality/quantity for producers, improve livestock production, incentivize hunting of invasive ungulates, and improve public relations, marketing, and conservation values for producers (Sergio et al. 2006, Ribaudo et al. 2010, Wilcox & Giuliano 2011, Brondum et al. 2017). Ecological benefits include better biodiversity conservation and decreased soil erosion in watersheds and coastal ecosystems. However, we note that the recovery of vegetation and soil is dependent upon how degraded the system became prior to the reduction in wild ungulate grazing, as well as factors outside of producers' control such as the ongoing drought conditions and upstream management of water and vegetation. Monitoring of these systems over the coming years, following the completion of fence building and expansion of control activities, will be critical to determine appropriate levels of wild ungulates within different fencing units. While the appropriate number in many units will be zero given the sensitivity of crops or ecosystems, a desire for commercial and community hunting opportunities may result in some non-zero number. Future studies are critical to support producers and land managers in determining what these numbers are and how to maintain them. Further, we highlight that while considerable progress has been made within the county of Maui Nui where Axis deer occur and impacts are dire, Mouflon sheep on the island of Hawai'i have not yet received equivalent efforts. It is our hope that the collaborative efforts and investments in Maui County will serve as a model for other islands experiencing similar impacts from other wild ungulate species.

Extension activities included publications, webinars, workshops, producer events, targeted youth outreach, and website development. To provide structured recommendations to producers based on our research results, we considered criteria from multiple stakeholders to determine IPM recommendations, seeking conservation of native species and culture while providing local food security through protection of crops, increased accessibility of protein from ranching and hunting, and decreased sedimentation run-off to coral reef fisheries. Results have been shared with communities in workshops with ranchers, producers, and other community members, annually at the Hawai'i Cattlemen's Convention both via tabling and in formal presentations (~500 attendees each year), at the Hawai'i Conservation Conference (~1200 attendees each year), in online public presentations, as well as via ongoing meetings with collaborative partners at ranches and government agencies. Two peer-reviewed publications regarding the direct and indirect economic impacts to ranches have been produced, one of which was published in 2024 and the other is currently under review. A two-chapter Master's thesis regarding forage impacts was defended this spring and will be available in May 2025. Video-based interviews with cultural practitioners regarding wild ungulates were obtained in spring 2025 and a synthesis of knowledge from these interviews will be complete in the summer of 2025.  Table1_ungulates_WSARE

Project Objectives:

Research Objectives:

  1. Quantify economic impacts of invasive, unmanaged ungulates to agriculture in the Hawaiian Islands. (completed fall 2024)
  2. Quantify impacts of unmanaged ungulates on forage production in agricultural landscapes in Hawai‘i. (completed spring 2025)
  3. Identify cultural significance of managed and unmanaged ungulates in agricultural landscapes of Hawai’i. (in progress, expected completion July 2025)
  4. Identify culturally appropriate and economically viable methods of control for unmanaged ungulates, in order to improve food security in Hawai’i. (completed spring 2025)
  5. Educate stakeholders and policy decision-makers about the impacts of unmanaged ungulates on economics, agriculture, food security, and ecosystems in Hawai'i. (completed fall 2024, but also ongoing)
  6. Develop decision support guidelines that facilitate stakeholder choices for economically viable and culturally appropriate Integrated Pest Management Practices (IPM) for these ungulate populations, in order to mitigate damage to agricultural lands and ecosystems in Hawai‘i. (in progress, expected completion July 2025)

Education Objectives

  1. Educate stakeholders and policy decision-makers on the impacts of wild ungulates on agriculture and food security through Peer-to Peer learning modules and workshops.

  2. Use Peer-to-Peer learning modules to increase awareness, and knowledge, skills, and abilities of stakeholders to recognize and discuss potential impacts of wild ungulates.

  3. Educate stakeholders on culturally appropriate IPM practices developed to control wild ungulate populations and mitigate damage to agricultural lands.

 

Timeline:

 

Year 1:

  • Stakeholder meetings/workshops to create initial framework (complete fall 2021)
  • Introductory Extension newsletter article (complete fall 2022)
  • Development and distribution of producer economic survey (complete winter 2022)
  • Initial project kick-off workshops, Peer-to-Peer Learning Modules and site visits with site partners, Co-PIs, extension agents, students and other stakeholders (complete spring 2023)
  • Field data collection for wild ungulate impacts to forage and crop production (complete summer 2023)
  • Cultural survey/interviews with ranchers, hunters, fishers, and other cultural practitioners (complete 2024)
  • Collection of producer economic survey data (complete summer 2023)

Year 2:

  • Compilation and analysis of cultural survey results and interviews (will be complete summer 2025)
  • Data compilation and analysis from producer economic survey results; factsheet produced (complete fall 2023, with 2nd paper complete 2024)
  • Data compilation and analysis from ecological field survey (complete spring 2025)
  • Build wild ungulate community science webpage (will be completed summer 2025)
  • Stakeholder meetings/workshops and Peer-to-Peer learning modules to discuss initial findings and next steps (complete 2024)
  • Construction of first draft of IPM decision guidelines (complete 2024)
  • Workshops, Peer-to-Peer learning modules and site visits with site partners, Co-PIs, extension agents, students, and other stakeholders  during IPM guidelines development, in order to optimize the utility of the guidelines (complete summer 2024)
  • Choose food security focus group (advisory team) members for each island (summer 2024) - check with Mark

Year 3:

  • Meet with food security focus groups to discuss island-specific issues (winter 2024) - check with Mark
  • Integration of all user feedback from workshops, Peer-to-peer learning modules and site visits with site partners, Co-PIs, extension agents, students, and other stakeholders to optimize the adoption of IPM decision guidelines (to be completed summer 2025).
  • Publications, reports, and extension materials prepared, submitted for review, and shared with stakeholders (to be completed summer 2025)
  • Evaluation of project outcomes and impacts using post project/activity evaluation and surveys on adoption of project recommendations (to be completed summer 2025)

Cooperators

Click linked name(s) to expand/collapse or show everyone's info
  • Lani Cran Petrie - Technical Advisor - Producer
  • Greg Friel - Technical Advisor - Producer
  • Jordan Jokiel - Technical Advisor - Producer
  • Kristin Mack Almasin - Producer
  • Jake Muise - Technical Advisor
  • Darren Strand - Technical Advisor - Producer
  • Carolyn Auweloa - Technical Advisor - Producer (Educator and Researcher)

Research

Materials and methods:

 

Methods (Organized by Project Objectives)

1.  Quantify the economic impacts of invasive, unmanaged ungulates to agriculture in Hawai

Survey design. A survey was designed to allow estimation of the value of wild ungulate damage and control for livestock producers in the Hawaiian Islands, including the severity of damage and damage costs, control costs, and descriptive data. The University of Hawai'i-Manoa Institutional Review Board (IRB) reviewed all survey materials and approved the survey on 26 September 2022 (IRB #2021-00877). Questions were based on previously deployed surveys in multiple geographic regions, including Anderson et al. (2019) and McKee et al., (2020) and were adapted to be appropriate to the context of the Hawaiian Islands and the goals of the present study.

Data were collected through a self-administered questionnaire hosted on Qualtrics (Qualtrics XM, Provo, UT, USA), an online survey platform, by the University of Hawai'i at Mānoa. A judgment sampling method was employed (Etikan and Bala, 2017). Judgment sampling is a non-random sampling method in which the sample is selected based on the researcher's knowledge of the units to be sampled. For this sample, the elements of the population selected for inclusion were known livestock producers on the islands. This method is used when a standard sampling method cannot be applied or is not appropriate given the aims of the study. As there is currently no known email (listserv), telephone, or mail  distribution list for the livestock producers on the islands, we identified representative subsets of the larger population through groups such as the Hawaiian Cattlemen's Council listserv (email distribution list),the Hawai'i Range and Livestock Management News newsletter listserv, and the Hawai'i Farm Bureau listserv.

The survey instrument was designed to elicit a range of values associated with wild ungulate presence. Although tailored in part to specifics of livestock production, the survey also collected information on property damage, control costs, and hunting practices. In addition to questions regarding potential disease spread from wild ungulates and related concerns, as well as pasture damage, producers were asked about livestock loss from depredation, disease, and other causes on their facility. Additionally, they were asked to report the costs of medical treatments and veterinary services related to wild ungulate contact with their livestock. Respondents first indicated their geographic location, which resulted in the electronic survey subsequently restricting which wild or feral ungulate species were included as potential responses to minimize inadvertently inaccurate responses as the presence of species varies among islands.

There were three core types of information collected by the survey: information on wild ungulate presence, damage, and control methods. The first type of information was the presence and recent changes in the population of wild ungulates in the producer's county and on their operation (Fig. 2, Shwiff et al. 2024). Wild ungulate presence provides a general indication of the economic threat they pose in the area through either direct damage or the risk of disease transmission.

The second core type of information collected was on wild ungulate damage to property (e.g.,  buildings, vehicles, equipment, roads, fecal contamination of resources), livestock losses, disease implications, veterinary care, medical costs, and pasture related damage. Property damage costs were assessed by asking producers to report damage due to destruction of structures, impacts on water resources used by livestock, and damage to pastures, trees, habitat, and other damage (Fig. 3, Shwiff et al. 2024).

Sections on livestock losses, disease transmission risk, and veterinary and medical costs followed (Fig. 4, Shwiff et al. 2024). Livestock medical and veterinary care was calculated as a sum of predation by wild pigs, disease, and medical and veterinary costs, which were all measured from value lost/costs associated with their highest and second-highest value livestock. As such, these do not capture the total costs of all their livestock, only their livestock with the highest value of production.

We asked producers to report information from 2021 because that was the most recent complete production year at the time that the survey was administered and was arguably easy for producers to self-report. Despite the potential inaccuracies with relying on self-reported damages, we chose this design for several reasons. First, self-reporting of wildlife damage to agriculture is common and has been show to be accurate way to reflect actual damage suffered by producers (Conover 2002, Johnson-Nistler et al. 2005, Tzilkowski et al. 2002, Wywialowski 1994). Second, livestock values can vary substantially according to the region and type of livestock (even with specific categories). We believed it preferable to rely on Hawaiian producers who have knowledge of local values and prices rather than making inferences from broader pricing statistics. Finally, even if producer perceptions vary from reality, producers often make decisions based on their perception of reality.

Control costs were calculated by summing all forms of control methods utilized by producers, including annual fence installation costs (Fig. 6, Shwiff et al 2024). The annual fence cost was the total cost to install a non-electric fence divided by the estimated years of use, provided by the producers. For the survey acres covered (39%), we determined a weighted average of $4.81/acre in damage plus control costs. The total amount of pasture acreage across all the islands is approximately 765,579 (Hawai'i Department of Agriculture, https://hdoa.hawaii.gov/salub/).

To estimate indirect costs associated with damage from wild ungulates on producers we focused on survey questions related to livestock, total herd values, and changes in composition or size of a producer's herd due to ungulate damage to pastureland. Specifically, questions 3.7 and 3.7 (Figure 2, Koppes et al. Under Review) assessed whether a producer had to reduce herd size due to loss of forage from ungulate damage and by how much they had to reduce their herd.

To estimate how much livestock producers were reducing their herd size, the midpoint of the range answered was used to determine the average percentage of herd reduction. A breakdown of this midpoint estimate method can be seen in Table 1 (Koppes et al. Under Review). Calculations were only applied to producers who reported their highest value livestock as cattle. Although producers were asked about several species such as sheep, goats, and other animals on their property, we only utilized estimates of cattle herd sizes provided by respondents, as, amongst all surveyed producers, the main livestock of concern was beef cattle with the average number reported by responded being 877 (range 0 – 15, 125; SD = 2,442). If the respondent answered question 3.7 but not 3.7a or answered ‘Don’t know’ to question 3.7a, their reduction amount was set to zero and therefore not included in the estimate of livestock loss.

This average reduction value was then used to calculate the original herd size using

equation (1) where 𝐻 is the original herd size of producer i, 𝐶 is the current herd size of producer

i, and 𝑟 is the percent that producer i reduced their herd in the last five years due to forage loss.

1) 𝐻𝑖 = 𝐶𝑖/(1−𝑟𝑖)

Livestock loss (or production loss due to herd reduction) was then calculated with

equation (2) where the livestock loss, 𝐿, for producer i is equal to the difference between the

original estimated herd size and the current herd size.

2) 𝐿𝑖 = (𝐻𝑖 − 𝐶𝑖)

The gross income lost for livestock producers in our dataset was calculated using

equation (3) where the livestock loss for producer i was multiplied by the producer’s average

price per head of cattle. Data on the per head price of livestock was acquired through self-reported livestock values in the survey. Since livestock values can vary substantially according to

region and type of livestock (even within specific categories), we believed it preferable to rely on

producers who have knowledge of local values and prices rather than making inferences from

broader pricing statistics. Additionally, even if producers are not entirely accurate in their perceptions of their livestock’s value, production decisions are largely driven by these

perceptions.

3) 𝐺𝑟𝑜𝑠𝑠 𝐼𝑛𝑐𝑜𝑚𝑒 𝐿𝑜𝑠𝑠 = Σ 𝐿𝑖𝑃𝑖

As cattle producers in the Hawaiian Islands usually range many females and only need one or very few males, when a producers reduces herd size, they would be losing birthing capacity. Additionally, we can reasonably expect this reduction in herd size to persist as, unlike wild ungulates in the contiguous United States, wild ungulates in the Hawaiian Islands are limited in their movements; therefore, their damage is likely to remain consistent. Based on these assumptions, our estimate of total income loss reflects the annualized cost of lowered production potential due to forage loss from wild ungulates occurring on cattle livestock operations.

2. Quantify impacts of unmanaged ungulates on forage production in agricultural landscapes in Hawai‘i.

Study Area. A variety of ungulates have been introduced across the Hawaiian Islands, with multiple species present on the two islands included in this study: Hawaiʻi and Maui. On the island of Hawaiʻi, wild pigs (puaʻa; Sus scrofa), mouflon sheep (Ovis musimon), feral sheep (Ovis aries), and feral goats (Capra hircus) have naturalized populations (Duffy & Lepczyk, 2021). The island of Maui hosts populations of wild pigs, axis deer (Axis axis), and feral goats (Figure 1). Wild pigs are commonly found in multiple ecosystems and are drawn to areas with supplemental food resources or fruiting trees, and are generalist feeders (Eisenberg 1981; Nogueira-Filho et al. 2009; Risch et al. 2022). On the island of Hawaiʻi, wild pigs are commonly found in native forests and grasslands (Nogueira-Filho et al., 2009; Stone, 1985); on Maui, they are concentrated in mid- to high-elevation forested areas (Risch et al., 2020). In grassland regions of Hawaiʻi Volcanoes National Park, grasses accounted for 50% of their diet (Baker, 1975). Feral goats are grazers and browsers depending on the availability of forage and are highly adaptable to their environment (Chynoweth et al., 2013). On both islands, feral goats are often found in dry lowland areas that are sparsely vegetated and range from coastal to sub-alpine areas (Chynoweth et al., 2013; Risch et al., 2022). On the island of Hawaiʻi, mouflon sheep are concentrated around Mauna Kea and extend into the northern slopes of Mauna Loa (Ikagawa 2013, 2013b; Judge et al., 2017a). Ikagawa (2013) estimated that mouflon occupy an area of 3,640 km², ranging from 550 meters in elevation to the summit of Mauna Kea (~3,660 meters). Axis deer on Maui are found in mid- to low-elevation shrublands and agricultural lands, with their range often overlapping with ranchlands (Risch et al., 2020). A survey conducted between late 2021 and early 2022 estimated a population of 46,743 axis deer, occupying 32% by area and occurring up to an elevation of 2,150 meters (Hess et al., 2022).     

Site locations. Four ranches agreed to serve as study sites, two on the island of Hawaiʻi and two on Maui, representing 47% of the total ranching area on the island of Hawai‘i and 42% of the total ranching area on Maui (Perroy & Collier, 2020). For the purposes of confidentiality, each participating ranch has been anonymized and assigned a unique letter identifier. On the island of Hawaiʻi, Ranch A is located on the southeastern side of the island. Ranch A consists of 29,000 acres of ranch land from sea level up to 5,000 ft and is part of a Cooperative Game Management Area (CGMA) which is a lease agreement with the State of Hawaiʻi Department of Land and Natural Resources (DLNR). Ranch A receives around 1,500 mm of annual rainfall. Ranch B is located in Waimea on the northern region of the island of Hawaiʻi. Ranch B consists of around 230,000 acres spanning from 2,000 to 8,000 feet above sea level and receives around 760 mm of annual rainfall. On Maui, Ranch C is located upcountry on the leeward slope of the Haleakalā volcano. This ranch consists of around 30,000 acres of land that range from sea level up to around 4,000 ft in elevation and annual rainfall spans from 430 mm to 4,750 mm. Ranch D is also located on the leeward slopes of Haleakalā. Ranch D contains around 18,000 acres of land, some of which are leased from the State of Hawaiʻi and private owners. Ranch D has an average annual rainfall of 890 mm.

Site selection. Survey locations were selected in coordination with the ranch managers in areas where managed livestock had been rotated out or were absent and where wild ungulates had been previously sighted. Survey points were then randomly selected within the parcel at a minimum of 100 meters apart. Due to accessibility considerations, points were in proximity to access roads. Research equipment (described below) was in the field for a minimum duration of thirteen days to minimize interference with ranch rotational grazing operations.

Ungulate detections. The following methods to capture wild ungulates relative abundance index (RAI) were derived from Risch et al. (2022). Each experimental plot consisted of six motion-activated game cameras (Bushnell Trophy Cams, Bushnell, Overland Park, KS) to detect wild ungulate activity. Cameras were arranged in a rectangular grid (two lines of three cameras), spaced 50 meters apart (Figure 2). Cameras were mounted onto nearby tree trunks when available or attached to t-posts anchored in the ground. To maximize the likelihood of detecting ungulates when they were in the area, cameras were angled toward areas with visible trails, tracks, or signs of damage. Cameras were set to take two consecutive photos after being triggered and then reset after three seconds.

Photos were organized by ranch site, point, and camera. The object detection model MegaDetector (Microsoft, 2020) was used in conjunction with Timelapse (https://saul.cpsc.ucalgary.ca/timelapse/) to identify photos with animals (Celis et al., 2024; Fennell et al., 2022; Leorna & Brinkman, 2022). Photo-viewing software Timelapse was used in conjunction with MegaDetector to manually classify animals by species. In Timelapse, an episode is defined as a sequence of images captured within a specified time span, representing an event such as animal movement through the area. For this study, episodes were generated using a one-minute time threshold. The total number of individual animals within each episode was recorded and used to quantify relative abundance metrics. The highest number of individuals recorded within an episode was divided by the number of camera trap nights to calculate the relative abundance index (RAI) (O’Brien et al., 2003; Palmer et al., 2018).

Grazing/biomass. Vegetation utilization by wild ungulates was quantified by comparing grazed areas (outside exclusion cages) to ungrazed areas (inside exclusion cages) (Bonham et al., 2004; Klingman et al., 1943). Three individual one m² grazing exclusion cages were randomly placed along three transects: two outside the camera array and one within the array, at the start of each monitoring period. These exclusion cages remained in place for the duration of the game camera monitoring. Rainfall at each point was measured using rain gauges placed at one of the point's exclusion cages and measured at the end of the monitoring period.

Forage biomass samples were collected from inside and outside the exclusion cages following methods adapted from Thorne and Stevenson (2007). A 0.22 m² (1.67 m circumference) sampling hoop was randomly placed near each exclusion cage to collect samples from outside exclusion cages, and samples from inside exclusion cages were collected from the center of the exclusion cage. Both the exclusion cages and sampling hoops were positioned in areas with grassier vegetation or in areas representative of the predominant vegetation. Vegetation biomass was measured by clipping live vegetation at ground level within the hoop and weighing it in a paper bag using a spring scale to determine the mass of fresh vegetation. One sample was collected at the beginning of the survey period prior to placing cages, and at the end of the survey period three samples were collected from inside the exclusion cages and three from outside. To determine dry weight values, the vegetation samples were oven-dried at 65℃ for three days and then reweighed at facilities on each respective island. The dry weight measurements were converted to pounds per acre using hoop size conversion (dry weight grams x 40 = lbs/acre), and then to kilograms per hectare to estimate the average available forage production. Wild ungulate grazing pressure was calculated by determining the average difference in forage biomass (kg/hectare) between the inside and outside samples of each grazing exclusion cage at each point. All values were averaged at each point. The initial samples collected were compared to samples at the end of the survey to ensure the forage collection was within a reasonable range for data integrity.

Plant Cover. At the start of the ungulate monitoring period, the line-point intercept method was conducted to assess ground cover. Three 100-meter transects were placed on the outsides and middle of the game camera array using a tape measure and at every meter, a pin was utilized to record the species at each intercept (100 points per transect) (Bonham et al., 2004; Elzinga et al., 1999). At every point, the number of detections of each species on a transect was averaged and converted into a percent cover and percent composition of foliar cover by species. Each plant species was categorized by grass, forb, shrub, litter, or bare ground. Litter refers to dried and dead plant material, while bare ground is defined as any surface lacking plant cover, such as exposed soil or rock. Species were identified in the field using iNaturalist and confirmed using the Hawaiʻi Range & Pasture Field Identification Guide and Weed’s of Hawaiʻi’s Pastures and Natural Areas when possible (CTAHR: Weeds of Hawaii’s Pastures and Natural Areas; An Identification and Management Guide, n.d.; Hawaii/Pacific Islands Area | Field Office Technical Guide | NRCS - USDA, n.d.; iNaturalist, 2024). For each transect, the elevation and compass direction were recorded.

Analysis. As ungulate community composition varies by island and location (Duffy & Lepczyk, 2021; Tomich, 1986), we grouped ungulates for analysis by either “all ungulates” or “grazing ungulates” for subsequent analyses; where the number of detections of an ungulate species was adequate, we ran individual models for each species.

The following analyses were conducted in R 4.1.1 (R Core Development Team, 2021). A linear mixed-effects model was implemented using the lme4 package to assess the relationship between forage samples inside the exclusion cage (kg/ha) and the fixed effects of rainfall and elevation, with site included as a random effect. To identify the best-fit model among all possible combinations, the MuMIn package was used to rank models based on Akaike’s Information Criterion (AIC) (Bartoń, 2023). The same approach was applied to examine the relationship between forage samples inside and outside the exclusion cage (kg/ha), with fixed effects of wild grazing ungulates RAI (feral goats, feral sheep, axis deer, and mouflon sheep), rainfall, and elevation, and site as a random effect. Forage samples inside the exclusion cage were assigned an RAI of zero, as they represent forage availability without exposure to wild ungulates.

To further assess wild ungulate relationships with forage availability, a linear regression analysis was conducted using the stats package to examine the relationship between average forage biomass loss (kg/ha) and RAI, including all wild ungulate species detected across islands (wild pigs, feral goats, axis deer, feral sheep, and mouflon sheep).

To evaluate the relationship between grazing wild ungulates RAI and percent cover of grass, forb, shrub, litter, bare ground, and rainfall, a Generalized Linear Model (GLM) framework was used. Model selection was performed using the glmulti package which automated model selection based on AIC values and Akaike’s weights (Calcagno & de Mazancourt, 2010)

3. Identify cultural significance of managed and unmanaged ungulates in agricultural landscapes of Hawai‘i.

To explore the cultural and experiential dimensions of wild ungulate management in Hawaiʻi, we conducted a series of semi-structured interviews with individuals engaged in land-based practices and natural resource stewardship. This qualitative approach aimed to capture the lived experiences, knowledge systems, and values that inform attitudes toward ungulate control, and to provide narrative depth to the broader survey and ecological findings of this study.

Participants were selected through purposive and snowball sampling strategies. Our project collaborator, Carolyn Wong, utilized her extensive network within the agricultural and resource management communities to identify and interview participants with firsthand experience in ranching, farming, hunting, wild ungulate harvesting, and cultural stewardship. Additional interviewees were identified through referrals from initial participants, allowing us to include a diverse range of voices and perspectives across the islands. This iterative sampling process continued until we reached information saturation, or the point at which no new themes were emerging within stakeholder groups.

All interviews were conducted in-person between late 2023 and early 2024. The semi-structured format allowed for flexibility in conversation while ensuring that key topics were consistently addressed across interviews. These included: unmanaged ungulate species of concern; managed livestock and their cultural significance; impacts on native species and ecosystems; paniolo (ranching) traditions; hunting and fishing culture; and food-related cultural practices. Interview questions were developed in consultation with previous cultural and social science projects in Hawaiʻi (e.g., Luat-Hūʻeu et al., 2023; Sato et al., 2018) and designed to elicit both individual experiences and broader cultural perspectives on land use and wildlife.

In total, 18 individuals participated in these interviews, generating approximately 244 minutes (just over four hours) of recorded dialogue. All interviews were digitally documented by team member Daniel Emhof. Transcripts are currently being reviewed for emergent themes related to culturally appropriate and economically viable wild ungulate control strategies. These interviews will be analyzed using a thematic coding framework to identify commonalities and differences in perspectives, with a particular focus on how cultural values and land-based knowledge can inform sustainable and culturally sensitive Integrated Pest Management (IPM) approaches.

4. Identify culturally appropriate and economically viable methods of control for unmanaged ungulates, in order to improve food security in Hawai‘i. 

We integrated both quantitative and qualitative data sources to examine wild ungulate control practices currently used by livestock producers across Hawaiʻi, and to assess their perceived effectiveness, economic feasibility, and cultural relevance. Specifically, we combined survey data from a statewide assessment of ranchers and livestock producers with insights gathered through semi-structured interviews with key land-based collaborators.

The survey instrument, administered between February and June 2023, included targeted questions about the types of control strategies producers currently employ—such as shooting on sight, hunting with or without dogs, trapping, and the installation of exclusion fencing (described in detail above). Respondents were also asked to evaluate the effectiveness and relative costs of each method. Because producers could report using multiple control approaches simultaneously, the survey captured nuanced patterns of management and provided a statewide snapshot of preferred and viable strategies under current economic conditions. These data were analyzed to identify which strategies were most frequently used, how producers evaluated their success, and what financial constraints or trade-offs were associated with their use.

To complement and contextualize these findings, semi-structured interviews were conducted with a diverse group of stakeholders including ranchers, farmers, hunters, commercial wild ungulate harvesters, and cultural practitioners. Interviewees were selected to reflect a range of perspectives and land-based knowledge systems across the islands. Interviews focused on individual experiences with ungulate control, cultural attitudes toward different species, and perceived barriers or opportunities related to management efforts. In particular, participants were asked to reflect on which control methods align with local values and cultural practices, and which approaches they believe are most sustainable for Hawaiʻi’s communities and ecosystems.

Together, the survey and interviews provide a holistic understanding of wild ungulate control in Hawaiʻi—linking economic viability with cultural appropriateness. These data will be analyzed both independently and in tandem to identify intersections where effective control strategies also align with local values and practices, providing a foundation for integrated pest management recommendations that are both actionable and place-based.

5. Educate stakeholders and policy decision-makers about the impacts of unmanaged ungulates on economics, agriculture, food security, and ecosystems in Hawai'i. This objective is integrated throughout all objectives, but is detailed under the Education Plan. 

6. Develop decision support guidelines that facilitate stakeholder choices for economically viable and culturally appropriate Integrated Pest Management Practices (IPM) for these ungulate populations, in order to mitigate damage to agricultural lands and ecosystems in the Hawaiian Islands. Island-specific, culturally-appropriate decision support guidelines, tailored to rancher and land manager values, were developed in consultation with producers and other stakeholders, in order to facilitate choices of IPM practices for their specific locations in regard to control of unmanaged ungulate populations (Years 2-3). Guidelines are specific to production goals, willingness to allow hunter access, location (due to correlated variables such as rainfall, temperature, wild ungulate type and abundance), as well as island-specific available actions such as hunting associations, county working groups and ordinances, and other geographically-specific information. Based on data collected during this project and user inputs, this project identified solutions that account for social, cultural, and economic variables. Outputs also direct the users toward island-specific contact information for permitting, state and federal offices that can provide specific types of support to achieve the identified optimal set of actions, hunting associations, and the state and county Game Management Advisory Commissions, which serve as contact points for hunter-focused management solutions.

 

Research results and discussion:

Results (Organized by Project Objectives):

1. Quantify economic impacts of invasive, unmanaged ungulates to agriculture in Hawaiʻi.

Results

Shwiff et al. 2024, Pest Management Science

Survey results. The survey was administered from February 2023 to June 2023. It received 113 responses, 68 of which were complete and met validation requirements. These validation requirements included surveys that could not be used consisted of not applicable (NA) as a response to all questions, respondents who did not reside in Hawaiʻi, or respondents provided that same value for every response. Given these considerations, out of the 68 responses 45 were usable from the targeted audience of livestock producers (Fig 7, Shwiff et al. 2024).

These 45 surveys were received from five islands, representing approximately 39% of all reported pasturelands in the Hawaiian Islands (Table 1, Shwiff et al. 2024). Surveyed acres covered a significant amount of the pastureland for the islands of Hawaiʻi (43%) and Maui (46%), but lower percentages for Oʻahu (18%), Kauaʻi (6%) and Molokaʻi (6%). The survey results were compared to total pastureland data as reported by the Hawaiʻi Department of Agriculture in the latest reporting year (Hawaiʻi Department of Agriculture, https://hdoa.

hawaii.gov/salub/). There were no participants from Niʻihau or Lānaʻi in our survey. Regarding pasture estimates, we assumed for this analysis that all of the land reported in the survey was designated as pastureland.

Most of the respondents reported that wild ungulate presence in their country and on their operation had increased over the last 3 years. On Hawaiʻi, 96% of respondents indicated that wild ungulate presence had increased in their county and on their operation. For Maui, 100% indicated an increase in their country and 89% on their operation.

For most of the respondents, the livestock concern was beef cattle. The average number of livestock reported by respondents was 877 (range 0 – 15, 125, standard deviation = 2442). Beef cattle, goats, and sheep were the top three livestock species raised by respondents, and on average respondents owned two livestock species. On Hawaiʻi most producers raised cattle, with goats ranking second in terms of overall number of animals on their facility. On Kauaʻi and Molokaʻi surveyed producers primarily raised cattle. On Maui, producers reported that cattle were the primary livestock produced, with sheep ranking second which was the exact opposite of Oʻahu.

Wild pigs (present on all islands except Lānaʻi) and axis deer (present only on Maui, Molokaʻi, and Lānaʻi) were reported to cause the most damage to property and operations (Table 2, Shwiff et al. 2024). Additionally, among livestock producers on the island of Hawaiʻi (n = 28), significant damage was reported from mouflon sheep (17% of respondents), feral sheep (14% of respondents), and feral goats (21% of respondents). Producers from Maui reported a high degree of damage by axis deer (70% of respondents) and feral goats (20% of respondents).

Direct impacts. On a scale of 1 (causing minimal damage) to 5 (causing the most damage) based on the damage to their operations, most respondents rated feral pigs and axis deer as causing damage at a scale of 4 or 5 (Figure 8, Swhiff et al. 2024), while feral sheep, goats, and mouflon sheep were rated lower. Of the islands from which responses were received, axis deer are only present on Maui and Molokaʻi, and feral sheep, mouflon sheep, and hybridized feral and mouflon sheep are only present on Hawaiʻi island. The total annual cost of wild ungulates summed across all ranch operations represented in the survey was reported to be $1.42 million, with most of the costs resulting from direct damage and control costs (Table 3, Shwiff et al. 2024).

Additionally, livestock producers spent around $2.03 million in fence installation costs. The largest costs were from property damage, control, and pasture repoair (Table 3, Shwiff et al. 2024), but respondents were only asked to consider the two highest value livestock species, which were less than all livestock species raised for nearly all the producers. The summation of damage costs by island provides the opportunity to determine damage costs per acre by island. Combining the damage costs per acre with total acres allows for the extrapolation of these costs for the entire state. Two extrapolations were made. First, we utilized the damage cost per acre for the total number of surveyed acres ($4.80) and multiplied this by the total number of pastureland acres in Hawaiʻi (756,579) to determine an annual estimate of over $3.6 million in damage and control costs to Hawaiian livestock producers annually. Second, we utilized the calculated damage cost per acre for each specific island and then extrapolated that value across only that island and then summed across the islands. Extrapolating costs this way led to an overall annual cost to the state of over $7.5 million.

Producers could utilize multiple methods of control so percentage will not add to 100%. The largest cost of control, other than installation of non-electric fences, was reported from shooting wild ungulates. Excluding fencing, a variety of control methods were used, including (listed in order of respondent preference) shooting on sight, hunting with dogs, hunting without dogs, and trapping, but no respondents used repellent. Examining this same information by island reveals that producers prefer control methods suitable for the type of invasive ungulates that occur on that island (Figure 9, Shwiff et al. 2024).

To a lesser extent producer reported that trapping was effective, and the use of repellents was viewed as not effective. Respondents reported that shooting and hunting with dogs were the most effective control methods with respect to reducing wild ungulate damage (Figure 10, Shwiff et al. 2024).

Koppes et al (Under Review), Rangeland Ecology and Management

Indirect impacts. Approximately, 35% of survey respondents indicated reducing their herd size due to wild ungulate damage on rangeland. Of these producers, 76% responded that the reduction impacted their breeding herd, consistent with our previous assumptions that herd reductions diminish birthing capacity. The producers who experience a reduction in the carrying capacity of their rangeland account for approximately 16% of the total acres surveyed and over 96% of all reported damaged acreage. Among these respondents, 6% reported not knowing by how much they’ve reduced their herd, and loss values were subsequently not calculated; 41% reported reducing between 1 to 25%, 35% reported reducing between 26 to 50%, 12% reported reducing between 51 to 75%, and approximately 6% reported reducing their herd size between 76 to 100% (Figure 5, Koppes et al. Under Review).

While wild ungulate damage was reported on the islands of Hawaiʻi, Maui, Molokaʻi, and Oʻahu, reduction of livestock cattle due to forage loss was reported on only three of these islands (Hawaiʻi, Maui, and Molokaʻi). Livestock producers on Maui account for roughly 81% of the reduction in production potential while Hawaiʻi and Maui account for 17% and 2% respectively. The total cost of livestock loss due to forage loss estimated from survey reports is $2.31 million (Table 2, Koppes et al. Under Review). Summation of livestock losses by island provides the opportunity to determine income loss per acre by island. Combining the loss per acre ($7.96) with total acres (705,199) allows for extrapolation of these costs for the entire state. We limited our extrapolation to three islands (Hawaiʻi, Maui, and Molokaʻi) as these were the only islands where ranchers reported herd reductions. Using the extrapolation above, we estimated statewide economic loss of $5.6 million due to herd reduction.

Producers who had to reduce their herd due to forage loss seemed to be, on average, 15 years younger, and the respondents were more often women. Those reducing herd sizes had larger increases in wild ungulates on their ranches, higher supplemental feed costs, a higher concern regarding parasites, and had substantial amount of damage, erosion, and acreage loss (Table 3, Koppes et al. Under Review). They also differed in their management actions to address these impacts and concerns. Those who had greater reductions in herd size also hunted and culled more wild ungulates, more often used dogs to hunt pigs (Figure 6, Koppes et al. Under Review) and worked more closely with government agencies.

To rule out pasture size as a driver for these differences, we examined overall respondent reported acreage between the two groups (reduced vs. non-reduced). The average size of a non-reduced producer’s rangeland (11,061 acres; SD = 32,805) was 44% larger than that of a reduced producer’s rangeland (6,129 acres; SD = 10,804), so the differences in acreage alone did not drive the results.

When damage rates on a scale of 1 to 5 were broken down by species and reduction status, we observed that reported damage for most species (Black-tailed deer, feral goats, feral and mouflon sheep, and wild pigs) remained relatively consistent across the two groups. Where they differed dramatically was reported damage due to Axis deer (Figure 7, Koppes et al. Under Review), suggesting that the impacts are highest on islands with invasive Axis deer.

Discussion

Wild ungulates impose substantial economic costs on Hawaiʻi’s cattle industry, with damages totaling between $9.2 million to $13.1 million in combined direct and indirect costs annually (Shwiff et al. 2024 and Koppes et al. Under Review). These expenses represent a significant portion of the industry’s five-year average gross income of $53.1 million, highlighting the economic strain wild ungulates place on ranchers (State of Hawaii 2023). Despite recent gains in production and revenue, such high costs from wild ungulate damage threaten the industry’s financial stability and growth potential. Addressing these impacts is crucial for sustaining Hawaiʻi’s cattle ranching operations and mitigating further economic losses. The demographic differences between reduced and non-reduced producers also offers insights into potential management approaches. Younger producers who are more affected by these challenges are engaging actively in ungulate management but may have less established practices or resources. This trend points to a critical opportunity for targeted support and education, particularly for newer farmers who may lack knowledge or means to effectively mitigate ungulate damage.

Damage was not equally distributed among islands, a result that is not surprising given that most ranching operations are on the island of Hawaiʻi, the largest island in the Hawaiian Archipelago, and that wild ungulate assemblages differ by island. The total reported property damage costs were highest on the island of Hawaiʻi, which is consistent with the fact that most of the survey respondents were from this island and roughly 39% of the total pastureland was accounted for from the survey respondents. However, when considering the herd reductions as a result of wild ungulates, the impacts were disproportionately felt on Maui, where 81% of all producers who reported reducing herd sizes were located (Koppes et al. Under Review). A result seemingly heavily driven by the impacts of Axis deer.

Respondents on all islands reported moderate to high damage to livestock production from feral pigs, with the highest reports of damage from respondents on the island of Hawaiʻi. Producers tend to underestimate the costs associated with wild pig damage by a factor of three (Carlisle et al. 2021), indicating that combined direct and indirect costs of wild ungulates could potentially range from $25 to $40 million annually. Although the island wide extrapolated estimates from this study are significant, these annual costs likely underestimate the total cost impact to ranchers for several reasons. First, respondents were only asked to consider their two highest valued livestock enterprises, which were less than all livestock species raised for nearly all the producers. Second, the economic burden of wild ungulate damage to pasture and property on livestock operations is not limited to lost production or increased production costs; it also includes the substantial additional cost of control efforts and risk of disease transmission and less tangible ecological costs such as the value of lost soil, reduced soil health, and reduced hydrological function.

Many producers reported applying a suite of control methods, the most common being shooting wild ungulates on sight and hunting with and without dogs. Shooting on sight was also perceived as the most effective in comparison to the other methods, similar to previous studies (McKee et al. 2020). Trapping and the use of repellants were reported as less commonly used and were also perceived as less effective. Producers continue to employ methods such as trapping even though these methods are perceived as less effective, in part because these methods can be used concurrently with other methods such as shooting on sight and hunting because wild ungulates adapt to the use of any particular method over time, emphasizing the importance of not only using a suite of methods but developing new methods over time. More research on effective control methods or combinations of control methods could potentially benefit producers in their fight against wild ungulate damage. Furthermore, given that hunting pressure and the presence of hunting dogs may cause ungulate to change their distribution and behavior (Risch et al. 2022), and the need for increased fencing, collaboration among land managers is likely critical to succesful implementation of these methods.

Our research to quantify wild ungulate damage to the Hawaiian Islands suggests that improved ungulate control may play a crucial role in food safety and safeguarding livestock, as well as promoting a One Health approach to management that considers interconnections between humans, wildlife, and the environment. Targeted, well-informed, collaborative, and consistent management of invasive wild ungulates necessitates an understanding of their economic impact on livestock producers. By understanding this damage, it is posible to provide management solutions that are not only economically viable for the producer, but are considerate of the fragile island ecosystem.

2. Quantify impacts of unmanaged ungulates on forage production in agricultural landscape in Hawaiʻi.

Results

Surveys were conducted from June 2023 to March 2024. In 2023, a total of 17 sites were surveyed: 5 on Hawai'i Island and 12 on Maui. In 2024, 18 sites were surveyed: 10 on Hawai‘i Island and 8 on Maui. Three sites at Ranch D were retrieved after the 13-day period due to the catastrophic fires that began on Maui on August 8, 2023; these sites were not affected by the fires. Additionally, two study sites were established in fire-affected areas at Ranch C following a wildfire event.

On the island of Hawaiʻi where there are populations of wild pigs, mouflon sheep, feral sheep, and feral goats we detected all species except the feral sheep. Wild pigs were detected at 73% of the sites (n = 11), with 368 photos captured, 50 episodes recorded, and a maximum group size of 3 individuals. mouflon sheep were also detected at 73% of the sites (n = 11), with 11,190 photos, 1,516 episodes, and a maximum group size of 52 individuals, the highest recorded species detected on the island of Hawaiʻi. Feral goats were expected to be seen at 27% of sites (n = 4) sites based on distribution models but were only detected at 13% of sites (n = 2), with a total of 92 photos taken, 5 episodes, and a maximum group size of 15 individuals.

On Maui, all expected wild ungulates were detected including wild pigs, feral goats, and axis deer. Wild pigs were detected at 75% of the sites (n = 15), with 3,336 photos, 176 episodes, and a maximum group size of seven individuals. Feral goats were found at 5% (n = 1) of the sites consistent with expectations based on species distribution models, and producing 1,622 photos, 179 episodes, with a maximum group size of 6. Axis deer were detected at 95% of the sites (n = 19), generating a substantial 92,152 photos. There were 4,637 episodes recorded and the maximum group size for axis deer was 127 individuals, the highest recorded across all sites.

A linear mixed-effects model was fitted to assess the effects of elevation (m) and rainfall (mm) on forage production (kg/ha) for samples collected from inside exclusion cages (i.e., without exposure to wild ungulates), with site included as a random intercept. Model selection revealed that the top model included only elevation (m) as a fixed effect, with site as a random effect (Table 2; AICc = 1885.79 weight = 0.53), while models incorporating rainfall received less support (ΔAICc ≥ 1.91). The following linear mixed-effects model evaluated the effects of elevation (m), rainfall (mm), and wild ungulate abundance (RAIgrazing) on forage production (kg/ha) across all samples, including those from both inside and outside exclusion cages. Elevation was identified as the strongest predictor of forage production through model selection (Table 3; AICc = 3688.11, weight = 0.36). The second-best model, which included both elevation and RAIgrazing, received similar support (Table 3; ΔAICc = 0.51, weight = 0.28).

Linear regressions were utilized to assess relationships between average vegetation biomass (kg/ha) and wild ungulate RAI. Feral goats and feral sheep were not assessed individually as they occurred at none or at a small number of sites (Table 2.1). The relationship between wild ungulates RAI and the forage loss showed a slight positive trend (Figure 2.5; Intercept = 212.43, SE = 183.92; Slope = 17.79, SE = 15.49, t = 1.15, = 0.04). On the island of Hawaiʻi there was a negative trend between the total wild ungulates detected and forage loss (Figure 2.6; Intercept = 234.57, SE = 248.08; Slope = -50.44, SE = 30.47, t = -1.66, R2 = 0.17) and Maui had a positive relationship between the total ungulates detected and forage loss (Figure 2.6; Intercept = 446.18, SE = 235.34; Slope = 25.41, SE = 16.77, t = 1.52, R2 = 0.17). For mouflon sheep on the island of Hawaiʻi, there was a negative relationship between RAI and forage loss (Figure 2.6; Intercept = 352.35, SE = 376.84; Slope = -61.86, SE = 39.94, t = -1.55, R2 = 0.11). There was a positive relationship between the RAI of axis deer and forage loss (Figure 2.6; Intercept = 451.57, SE = 231.78; Slope = 25.72, SE = 16.64, t = 1.55, R2 = 0.11).

Plant species were organized by functional group (grass, forb, shrub, and lichen) see Table A1 Appendix A. The automated model selection using the glmulti function evaluated 32 models of all possible combinations of wild grazing ungulates relative abundance (RAIgrazing) with predictor variables: grass, forb, shrub, bare ground cover, and rainfall. The model with the lowest AIC value was the null model (RAIgrazing  ~ 1) with an AIC of 261.62 and 0.091 weight (Table 4), indicating that none of the predictor variables had a significant effect on the relative abundance of wild grazing ungulates across study sites (weights = 0.0005 - 0.091).

Discussion

            This is the first field study to examine the relationship between forage loss and wild invasive ungulates on ranchlands in the Hawaiian Islands. We found substantial vegetation loss at 66% of the study sites, consistent with substantial losses reported by ranches surveyed in this WSARE project. Further, wild ungulates were detected at sites where ongoing impacts from drought and wildfires resulted in a lack of forage availability for livestock, suggesting that wild ungulates may continue to degrade pastures long after they are unsuitable for ranch production.

Elevation emerged as the primary driver of forage production in both models, highlighting its strong influence on vegetation growth across sites (Table 2 & 3). The top model for forage production inside exclusion cages included only elevation as a fixed effect, suggesting that, in the absence of wild ungulates, elevation has a strong impact on forage availability (Carlyle et al., 2014; Wu et al., 2024; Zhang et al., 2023). This is likely due to environmental differences at varying elevations such as temperature (K. Yang et al., 2022; Y. Yang et al., 2023), soil fertility (Khosravi Aqdam et al., 2023), and vegetation structure (Bora et al., 2021; Han et al., 2022). The strong influence of elevation on forage production in this study aligns with findings from research on native plant cover and fire dynamics in Hawaiian ecosystems (D’Antonio et al., 2000). This study similarly identified elevation as the primary environmental driver of vegetation responses, whereas rainfall did not systematically explain variation in plant cover. When assessing forage production across all samples inside and outside exclusion cages, elevation remained the strongest predictor.

Wild ungulate abundance appeared in the second-best model suggesting that grazing pressure may also influence forage availability, although to a lesser extent than elevation. This result aligns with previous studies demonstrating that ungulate grazing reduces forage biomass alongside other environmental factors (Frank et al., 2016; Ibañez-Alvarez et al., 2022; K. Kramer et al., 2006). The relatively close model weights between the top two models indicate some uncertainty regarding the relative influence of grazing compared to elevation, suggesting further investigation into potential interactions between these factors. Rainfall, while often considered a key factor in plant productivity (Gibson-Forty et al., 2016; Heisler-White et al., 2009), did not significantly improve model performance (Table 2 & 3), but we note that there has been a multi-year drought which may have influenced study results at some sites due to severe soil alteration.

The sites where grazing ungulates were detected (n = 32), 66% had documented forage loss across the island of Hawaiʻi and Maui (Appendix Table 2). Losses per site ranged from 74.72 kg/ha to 1942.81 kg/ha in areas with preferred cattle forage. To put this into context, in two weeks ungulates consumed the food needed for 6 to 161 animal units (AU) (450 kg cow) for one day assuming one AU consumes 12 kg of dry matter per day (Heitschmidt & Stuth, 1991; Holechek, 1989; Scarnecchia, 1990; USDA, 2003). These results align with reports of ranchers reducing stocking rates or no longer rotating cattle through certain pastures due to impacts from wild invasive ungulates.

Multiple factors such as rainfall, drought, and soil health, interact with ungulate abundance to affect forage availability for ranching. Additionally, some sites were in states of regrowth following fire damage (Ranch C, sites 3 and 4), were dominated by non-preferred vegetation (Ranch D sites 4 and 7), or lacked vegetation (Ranch C, site 4; Ranch D, site 10), likely influencing visitation rates by wild invasive ungulates. Despite these conditions, wild ungulates were present, suggesting their ability to persist in areas unsuitable for cattle further exacerbating the continued alteration of pastureland.

Wild grazing ungulates (feral goats, axis deer and mouflon sheep), did not show a preference for specific vegetation types—such as grass, forbs, shrubs, or bare ground—and rainfall was not a significant predictor of their relative abundance. These findings suggest that these introduced wild grazing ungulates are opportunistic generalists, highly adaptive to varying environments (Barroso et al., 2000; Freschi et al., 2021; Newman et al., 1995; Rogosic et al., 2007; White et al., 2023). Wild ungulates may also be limited in their ability to move to areas with preferred vegetation due to barriers such as fencing or urban development. 24 out of the 35 sites had fireweed (Senecio madagascariensis) present which is a known problematic pasture weed (Haselwood et al. 1983, Le Roux 2010). The weeds compete with preferred grasses such as Kikuyu grass (Pennisetum clandestinum), which is an important pasture grass due to its adaptability (Fukumoto & Lee 2003). Due to wild ungulates adding unwanted grazing pressure on ranchlands, grasses become suppressed, allowing noxious weeds and toxic forbs to grow in place (Leopold & Hess 2016, Perkins et al. 2018).

While not significant, detections of axis deer and mouflon sheep had the highest correlation with forage loss on the islands where they occur. Previous studies have indicated that grazers reduce forage production and cause significant vegetation damage when overabundant or introduced (Marchiori et al., 2012). As our study found areas with forage loss but did not detect strong trends or significant relationships with the abundance of wild ungulates, this may suggest that vegetation damage is extensive and immediate control efforts should be implemented.

Study limitations included relatively small sample sizes for feral goats and feral sheep (Table 2.1). Additionally, the short data collection period (two weeks), limited by rotational grazing schedules, likely captured only a snapshot of the complex interactions between wild ungulates and vegetation, potentially missing seasonal variations in forage dynamics or wild ungulate behavior. Ranchers have observed ungulates moving away from certain areas after heavy rainfall (pers. Comm. Lani Petrie.). Future studies should address these limitations by increasing the number of study sites, extending data collection across multiple seasons, and integrating data such as GPS tracking of ungulate movement, vegetation recovery rates, and soil health metrics to provide a more comprehensive understanding of these interactions.

This study advances understanding of the complex relationships between invasive wild ungulates and forage loss, and points toward interactions among ecological factors beyond wild ungulate abundance alone. Further, these results highlight the need for targeted removal of wild ungulates to reduce impacts to local ranch production. Long-term research is crucial to capture the cumulative impacts of ungulates on forage loss and the environmental factors driving their movement, such as vegetation structure or rainfall patterns. Future studies should also explore ungulate impacts across diverse habitats, including native forests and agricultural crops, to develop effective management strategies for mitigating their widespread effects.

3. Identify cultural significance of managed and unmanaged ungulates in agricultural landscape of Hawaiʻi.

Results

Semi-structured interviews were conducted by our project collaborator, Carolyn Wong and digitally documented by Daniel Emhof. Interviews consisted of 18 individuals whose roles span a range of land-based practices and experiences, including ranching, farming, hunting, commercial wild ungulate harvesting, and cultural stewardship. Many participants identified with more than one role, reflecting the interconnected nature of these practices in Hawaiʻi. A total of 18 individuals were interviewed for a combined 244 minutes (4 hours) of interview time. All interview footage was collated by February, 2025 and we were in the early stages of reviewing these interviews for cultural and historical context but due to the stop work order issued by the Trump Administration work is just now resuming. For those reasons, results and conclusions will be included in the final report. We anticipate these interview videos will provide valuable insights in addition to the work already described above.

4. Identify culturally appropriate and economically viable methods of control for unmanaged ungulates, in order to improve food security in Hawaiʻi.

Results

Survey responses revealed that producers across the state rely on a suite of control methods to manage wild ungulate impacts. The most commonly reported approaches included shooting on sight, hunting with or without dogs, and fencing. Shooting on sight was the most widely utilized and was also ranked by producers as the most effective strategy. Hunting with dogs was also commonly employed and considered effective for targeting specific species like wild pigs. In contrast, methods such as trapping or the use of repellents were less frequently used and generally perceived as less effective.

In terms of economic feasibility, fencing represented the highest cost control method, with producers reporting over $2 million spent in recent years on non-electric fencing installations. These investments, while often necessary to protect pasture and infrastructure, present a significant financial burden—especially for newer or smaller operations. Producers commonly noted that no single method was sufficient on its own, and that wild ungulates often adapt to repeated use of one technique. As such, many ranchers reported rotating among several control strategies over time.

To complement these quantitative findings, semi-structured interviews were conducted with 18 individuals representing ranchers, farmers, hunters, cultural practitioners, and commercial harvesters. These interviews, totaling over four hours of recorded dialogue, are currently being reviewed and analyzed. Once completed, they are expected to provide critical insight into the cultural dimensions of ungulate control in Hawaiʻi—highlighting which strategies align with traditional values and community stewardship goals, and where management practices may benefit from cultural grounding. These perspectives will be vital to understanding how control efforts can be made more inclusive, effective, and sustainable in the long term.

5. Educate stakeholders and policy decision-makers about the impacts of unmanaged ungulates on economics, agriculture, food security, and ecosystems in Hawaiʻi. This objective is detailed under the Education Plan. 

6. Develop decision support guidelines that facilitate stakeholder choices for economically viable and culturally appropriate Integrated Pest Management Practices (IPM) for these ungulate populations, in order to mitigate damage to agricultural lands and ecosystems in the Hawaiian Islands.

This project represents the first integrated effort in Hawaiʻi to combine ecological monitoring, economic analysis, and producer-informed perspectives to develop place-based decision support for managing invasive wild ungulates. Our research documented widespread and substantial impacts of unmanaged ungulates on ranching operations across the state, with economic survey data revealing up to $13.1 million in combined direct and indirect losses annually. Concurrently, field-based monitoring demonstrated that 66% of surveyed sites showed measurable forage loss directly attributable to wild ungulate grazing, with some sites exhibiting vegetation losses equivalent to forage needed by up to 161 cattle in just two weeks. These combined findings confirm that wild ungulates are a major production-limiting factor on Hawaiʻi’s rangelands, requiring coordinated, multi-pronged management approaches to mitigate damage and support ranching viability.

While producers employ a variety of control methods, fencing, community hunting, and aerial shooting emerged as key strategies with the greatest perceived effectiveness—particularly for managing axis deer. Commercial harvesting, though currently limited to a few operators, has also shown considerable success in reducing populations and holds promise as a viable business model. However, conversations with ranchers, land managers, and preliminary analysis of interview data revealed important reservations about the long-term viability and social acceptability of these approaches.

Fencing, while widely used and often necessary in high-priority areas, raises concerns among stakeholders regarding land access, maintenance costs, and the potential for increasing ungulate pressure in adjacent unfenced areas. Many acknowledge that fencing is essential in some locations—such as around sensitive habitats or highly productive pastures—but caution that it is not a feasible or equitable solution for landscape-scale or island-wide management. Similarly, aerial shooting has proven to be an efficient population control tool when requested by landowners and implemented by state agencies, but skepticism remains—particularly within the broader community—regarding its perceived wastefulness and lack of transparency in carcass recovery or use. These concerns highlight the importance of balancing technical effectiveness with public trust, transparency, and community buy-in.

Commercial harvest has been met with cautious optimism; while it demonstrates a functional model for both population control and food security, there is concern that as ungulate populations decline in some areas, the economic sustainability of the harvest model may falter. Taken together, these perspectives reinforce the need for a diversified, adaptive IPM framework—one that considers the trade-offs, limitations, and cultural sensitivities associated with each control method.

Additionally, the distribution of impacts and access to resources remains uneven across islands, with Maui Nui experiencing greater investment and coordination compared to areas like Hawaiʻi Island, where Mouflon sheep continue to exert heavy pressure on pasturelands. These disparities underscore the need for adaptable, region-specific guidelines that consider local species assemblages, land tenure, and existing community networks. Importantly, many control strategies are shaped not only by economics or logistics but by cultural values, access rights, and trust in management institutions.

To better capture these dynamics, we conducted semi-structured interviews with 18 individuals spanning ranching, farming, hunting, wild harvesting, and cultural stewardship. These interviews are currently being transcribed and analyzed, and preliminary insights are already informing our understanding of stakeholder needs and values. Once complete, this qualitative data will be integrated with our ecological and economic findings to guide the development of culturally grounded, economically viable recommendations that reflect the complexity of Hawaiʻi’s working landscapes.

These findings have already supported the development of peer-reviewed publications, outreach materials, graduate student theses, producer workshops, presentations at major venues such as the Hawaiʻi Cattlemen’s Convention, Hawaiʻi Conservation Conference, Hawaiʻi Invasive Pest Conference. Continued stakeholder engagement, adaptive management, and long-term monitoring will be essential to provide continued support to the agricultural producers in Hawaiʻi. 

Participation Summary
70 Producers participating in research

Research Outcomes

Recommendations for sustainable agricultural production and future research:

Ranchers across the Hawaiian Islands are facing mounting economic strain due to the growing impacts of unmanaged wild ungulates. Our statewide survey found that producers collectively spent over $3.4 million annually on damage repair and control, with fencing and property damage being the largest direct costs. When extrapolated to the full extent of Hawaiʻi’s pasturelands, these losses could total as much as $13.1 million per year. Over one-third of ranchers reported reducing herd sizes—greatly impacting breeding capacity—due to ungulate-induced forage loss. These reductions threaten long-term production capacity and disproportionately impact younger and female producers, many of whom are already operating with fewer financial resources. On Maui, where axis deer impacts were especially severe, herd reductions were particularly common, representing a significant threat to the island’s ranching viability.

Field data collected from ranches on Hawaiʻi Island and Maui further confirm the scale of this problem. At 66% of surveyed sites, wild ungulates caused measurable forage loss, with some areas showing the equivalent of forage consumed by between 6 to 161 cattle in just two weeks. Axis deer and mouflon sheep were especially associated with severe vegetation losses. These impacts were observed even in drought-affected and post-wildfire landscapes where pasture was already stressed or recovering—highlighting the resilience and adaptability of these invasive ungulates, and the ongoing risk they pose even in marginal or degraded areas. Our study also found that ungulates were not selective in vegetation type, indicating that these species are highly opportunistic grazers capable of exploiting a range of environments and further suppressing pasture recovery.

To add further depth and community perspective to these findings, we also conducted a series of interviews with ranchers, farmers, hunters, commercial wild harvesters, and cultural practitioners across the islands. These interviews—currently being reviewed and analyzed—are expected to provide invaluable insight into the lived experiences, values, and management decisions of those most directly affected by wild ungulates. By pairing these narratives with empirical data, we aim to build a more comprehensive, culturally grounded understanding of the issue—one that informs practical, place-based solutions and supports the long-term resilience of Hawaiʻi’s working lands and communities.

10 New working collaborations

Education and Outreach

20 Consultations
5 Curricula, factsheets or educational tools
2 Journal articles
10 On-farm demonstrations
3 Published press articles, newsletters
10 Tours
12 Webinars / talks / presentations
70 Workshop field days

Participation Summary:

150 Farmers participated
150 Ag professionals participated
Education and outreach methods and analyses:

Educate stakeholders and policy decision-makers on the impacts of unmanaged ungulates on agriculture and food security in Hawaii through Peer-to-Peer learning modules and workshops.

Year 1:

(1) We produced an extension newsletter article on the socioeconomic impacts of unmanaged ungulates, distributed via the Cooperative Extension website.

(2) We contributed to two meetings: (1) the Cal-Pac Society for Rangeland Management Fall 2022 Meeting October 6-7, 2022 in Waimea, Hawai'i; and (2) the Hawai'i Cattlemen's Convention and Annual Meeting November 18-19, 2022 in Waikaloa, Hawai'i. At the Cal-Pac Society for Rangeland Management meeting we presented a 15-minute talk on the challenges associated with the rising populations of unmanaged ungulates and their impacts on agriculture and food security. Four of the team members, including two Co-PIs and two producers also sat on a 5-person panel regarding the impacts of wild unmanaged ungulates on ranches in the Hawaiian Islands, and took questions from audience participants. At the Hawai'i Cattlemen's Convention we hosted an informational table where we talked with cow-hands and ranch managers attending the meeting regarding impacts on their ranches, and increased awareness of the upcoming economics survey to improve response rates. Team members present that week also visited ungulate damage areas to help with full understanding of the problem. 

(3) We attended the Maui Rangeland Health Workshop, on July 19, 2023. There were approximately 30 participants at the workshop hosted by Maui Cattlemen’s Association. We met with producers, State and Federal managers, and researchers regarding axis deer issues impacting ecosystem services, soil health, watershed function, aquifers, and coral reefs, as well as management for resilience to climate change. 

Year 2:

(4) A presentation was made by Derek Risch at the Invasive Pest Conference in Honolulu, Oʻahu on August 9-10, 2023 entitled “Distribution and abundance of wild ungulates in Hawaiʻi highlights their diverse impacts” and co-authored by M Price and J Omick. This presentation included discussion of unmanaged ungulates in the Hawaiian Islands and approximately 100 wildlife professionals attended. 

(5) A presentation was given by Derek Risch at the Seminar Series in the Department of Natural Resources & Environmental Management at the University of Hawai‘i at Mānoa entitled “Wild Ungulates in Hawaiʻi: Distribution and Abundance” and co-authored by M Price and J Omick. Approximately 50 wildlife professionals, graduate students, and University faculty members attended. 

(6) We reached out to beef producers via the Hawai’i Cattlemen’s Convention and Annual Meeting in Waikoloa, Hawai’i, in October 2023, with the following presentation:

“Economic estimates of invasive wild ungulate damage to livestock producers in Hawai’i” by SA Shwiff, K Caires, G Friel, L Katayama, Z Munoz, MR Price (presenting), D Risch, M Shartaj, K Steensma, M Thorne, R Zifko.

We also staffed a table during the two days of that meeting in order to facilitate ongoing discussion with attending producers. Approximately 125 people attended the presentation and 215 stopped at the table during the meeting.

(7) A poster presentation was made at the Wildlife Society Western Section Conference in Sonoma, California, February 6-9, 2024, by Lauren Katayama. The poster was entitled “Wild Ungulate Impacts on Ranchlands in Hawai’i” and co-authored by D Risch, M Thorne, K Caires, G Friel, K Steensma, C Auweloa, J Omick, S Shwiff, M Price.  Approximately 100 attendees viewed the poster.

(8) A presentation was made by Stephanie Shwiff at the Vertebrate Pest Conference in Monterey, California, on March 11-14 2024, entitled “An Update on the Economics of Wild Pig Damage in the US” which included discussion of unmanaged ungulates in the Hawaiian Islands. Approximately 100 wildlife professionals attended this presentation.

Year 3:

(9) In May 2024, a presentation was made by Karen Steensma to the Trinity Western University Coral Reef Ecology course in Kona, HI entitled "History of human interactions with reef ecosystems: Hawaiʻi as a case study" in which the impacts of invasive ungulates on Hawaiian agroecosystems was highlighted, including erosion impacts of overgrazing and land disturbance on Hawaiian reefs. This was followed by a field excursion with the same class, to the Kawaihae area to view extensive erosion from feral goats and resulting sedimentation on the adjacent reef, led by Carolyn Auweloa. There were 25 biologists in attendance for both the presentation and the field examination. 

(10) On May 15, 2024, we (Melissa Price, Mark Thorne, Derek Risch, Lauren Katayama) hosted a workshop at Ulupalakua Ranch to share key findings from our project with the ranching community and interested members of the public. Presentations were given by Derek Risch and Lauren Katayama, covering topics such as the history, distribution, and impacts of introduced ungulates across the Hawaiian Islands. Following the presentations, we facilitated a group discussion to engage attendees in meaningful dialogue about the challenges posed by ungulates and to explore potential tools, strategies, and community-based approaches for addressing these impacts.

(11) On May 17, 2024, we (Melissa Price, Mark Thorne, Derek Risch, Lauren Katayama) hosted a workshop at the College of Tropical Agriculture and Human Resource's Kamuela Extension Office to share key findings from our project with the ranching community and interested members of the public. Presentations were given by Derek Risch and Lauren Katayama, covering topics such as the history, distribution, and impacts of introduced ungulates across the Hawaiian Islands. Following the presentations, we facilitated a group discussion to engage attendees in meaningful dialogue about the challenges posed by ungulates and to explore potential tools, strategies, and community-based approaches for addressing these impacts.

(12) In November 2024, we tabled at the Hawaiʻi Cattlemen's Convention in Waikoloa, HI sharing project results via handout materials, presentation, and conversations with hundreds of attendees. A formal presentation was made by Stephanie Shwiff summarizing results of our work to date, and offering suggested pathways forward based on results from the economics survey. 

 

Education and outreach results:

A total of 43 ranchers at the Hawai'i Cattlemen's Convention signed up to receive survey results and more information on our project. In-person visits to five ranches by the majority of the team took place in the first year, with additional visits by extension team members (see below). 

Co-PI Kyle Caires:

Reached out to these additional ranches who agreed to let us setup and monitor wildlife for project.
*Nobriga Ranch, Kahakuloa and Waihee (2 locations)
*Peahi Farms, Haiku
*Diamond B ranch (3 locations) - Kaupo, Kula, Haiku/ Huelo
*J2S Ranch, Haiku
*Souza ranch, Ulumalu/Haiku
*Haleakala Research Station, Piiholo/Olinda/Makawao
 
Participated in stakeholder interviews with Professor Kathleen Merrigan in the Swette Center for Sustainable Food Systems at Arizona State University. Impacts of Axis Deer in Hawai’i and potential management solutions research study.  July 2023
 
Member, and Chair, Kula Agriculture Park Committee (KAP), Maui County. Appointed by the Mayor.
Member, Axis Deer Task Force. University of Hawaii at Manoa representative on Maui County's Axis Deer Task Force. Chair of Research and Education Committee.
 
Presented impacts from Axis Deer Damage to crops and pasture production at two axis deer task force meetings. The committee will also hear from Susie Thieman, executive director of Lōkahi Pacific, on the assistance program for axis deer relief, and Dr. Kyle Caires, animal scientist at the University of Hawaiʻi, on the Kula Agriculture Park and axis deer mitigation.
 
Member, and Co-Chair, Axis Deer Impacts Committee, Maui County Farm Bureau
Member, and Co-Chair, Axis Deer Impacts Committee, Maui's Agricultural Working Group
 
 
Grant Reviewer,  Maui County Axis Deer Microgrant For Farmers and Ranchers, Lokahi Pacific $1.5 million program.
 
Provided testimony on the following Bills to the Hawai Farm Bureau Federation at their Annual Meeting in October 2022
Provided testimony on the following Bills to the Maui County Council members November 2022, aired on TV, Akakū Channel 53
Provided testimony on the following Bills to the Maui County Chamber of Commerce members December 2022
 
HB 606, HD1 RELATING TO MEAT PROCESSING. Requires the department of agriculture, in collaboration with other state agencies, to develop and implement a plan by 7/1/2025 to expand the meat processing capacity in the State to allow for meat from axis deer and other wild game to be processed for distribution by non-profit food distribution services.  Appropriates funds.  Effective 6/30/3000.  (HD1)
 
HB 609, HD2. Appropriates funds to the department of agriculture for its meat inspection program, including three full-time equivalent (3.0 FTE) inspectors to perform meat inspection services.  Effective 6/30/3000.  (HD2)
 
HB 1382, HD2 - RELATING TO MEAT DONATION. Provides for donation of wild game to under-resourced communities.  Establishes a task force for development and implementation to expand meat processing capacity in the State.  Appropriates funds.  Effective 6/30/3000.  (HD2)
 
Reached out to these food aggregators as partners to help gain support for the legislative bills to help with Axis deer issues.
* Feed My Sheep, Maui
* Maui Food Bank
 
Interviewed by Maui News, regarding axis deer problem in Maui County. Conservation officials, ranchers, urge action on deer issues. Conservation officials, ranchers, urge action on deer issues
 
Interviewed on KHON2 news regarding axis deer problem in Maui County, September 2022. Maui venison could soon stock local grocery stores, https://www.khon2.com/local-news/maui-venison-could-soon-be-sold-in-local-grocery-stores/. Dr. Caires presented information regarding mitigating the axis deer problem, and opportunities for venison in local markets. Aired on KHON2 News TV for several news cycles.
 
Interviewed on KHON2 news regarding axis deer problem in Maui County, August 2022. Axis deer threatening Maui's natural resources. https://www.khon2.com/local-news/axis-deer-threatening-mauis-natural-resources/Dr. Caires presented pasture grazing data demonstrating the actual impacts of axis deer overgrazing on (1) row crop production for farming and (2) forage growth and recovery on recovery for pangola grass pasture. Aired on KHON2 News TV for several news cycles. https://www.youtube.com/watch?v=5SX1CGbq2ZU
 
Provided testimony on the Maui County Council's, Infrastructure and Transportation Committee meeting on Oct. 31 at 1:30 p.m. regarding axis deer updates across Maui County.
 
 
Hosted attendees and provided a field day site for the Pacific Regional conference of the National Association of Conservation Districts (september 2022). NACD members and attendees took tours through areas like Haleakala Ranch and the University of Hawaii’s Tropical Agriculture and Human Resources experiment station in Piiholo, as well as heard updates from local farmers, the state Department of Land and Natural Resources and the Maui Axis Deer Task Force. Dr. Caires presented cover crop field trial and pasture sites demonstrating the actual impacts of axis deer overgrazing on forage growth and recovery, 80+ in attendance. Dr. Caires provided a followup presentation during the evening program held at the Oskie Rice Arena on Kaonoulu Ranch.  “Imua—Building Capacity with Partnerships”—is the theme of the conference, which covers the Southwest and Pacific regions with close to 175 groups in all. The Hawai‘i Association of Conservation Districts, representing 16 Soil and Water Conservation Districts across the state, will host the event.
 
Presented impacts from Axis Deer Damage to crops and pasture production at two axis deer task force meetings (Summer and Winter 2022). The committee will also hear from Susie Thieman, executive director of Lōkahi Pacific, on the assistance program for axis deer relief, and Dr. Kyle Caires, animal scientist at the University of Hawaiʻi, on the Kula Agriculture Park and axis deer mitigation.
 
Provided testimony to county officials in Mayors Office and Department of Finance regarding funding for a Mobile Slaughter Unit for Maui Ranchers/Farmers. (sept 20, 2022)
Maui County Budget Director, Michele Yoshimura, spearheaded effort to discuss acquiring a Mobile Slaughter Unit for Desmond Manaba to assist Maui farmers and ranchers with the slaughter and production of axis deer.
 
 
Hosted professional development training for (1) MEDB high school agricultural interns and (2) High School Agricultural Teachers from Maui, Molokai and Lanai, at the University of Hawaii’s CTAHR - Haleakala Research Station (August 2022). Ag teachers and their students learned hands-on about topics such as cover crops, no-till planting, diversifying existing pastures with legumes, and axis deer mitigation efforts important for prevent erosion, and building soil health in order to have a production agricultural enterprise. We helped teachers focus on methods to adopt some of these  best management practices in their own school gardens and livestock facilities.
 
Year 2: A total of at least 200 producers and industry professionals attended our presentation or stopped at our table during the 2023 Hawai’i Cattlemen’s meeting. Additionally, at least 150 wildlife professionals attended our presentations at the Invasive Pest Conference in Honolulu, HI and the Seminar Series at the University of Hawaiʻi. An additional 100 conservation professionals viewed the Wildlife Society poster in Sonoma, CA, raising broader awareness of the issue. Another 100 people attended our presentation at the Vertebrate Pest Conference in Monterey, CA.
5 Farmers intend/plan to change their practice(s)
5 Farmers changed or adopted a practice

Education and Outreach Outcomes

Recommendations for education and outreach:

(1) Based on the first year of talking with stakeholders at workshops and events, it is clear that there are diverse perspectives and values regarding the wild ungulates. Some cowhands prefer to maintain some wild ungulates on the ranches to hunt for food for their families and friends. On other ranches there is zero tolerance for some types of ungulates (e.g., pigs or axis deer). Thus, shared solutions across ranch boundaries are critical to achieving goals, since the intentions of neighbors may differ regarding wild ungulates. In our second and third year we are developing tools to address this challenge.

(2) Secondly, many ranches are already implementing actions to address impacts of unmanaged ungulates. Over $6 million has been spent by ranchers to modify fences to exclude Axis Deer in the last year. Smaller ranches may not have the funds available to invest in this infrastructure, but strategic fencing funded by the State or Federal government could support these smaller ranches. Our models developed over the next two years may help direct fencing funds towards areas where they will most benefit ranch production. 

(3) In the second year, Hawai’i beef producers responding to our economic survey, and those engaging directly at the HCC meeting and through other contact, have indicated that aside from exclusion fences, hunting access is a key issue. Some do their own hunting, some contract with hunters, and some allow other hunting access. However, open hunting access is not without risk and many ranchers are wary of that. Thus our development of decision-making tools over the next year will include separate workshops for ranchers, for hunters, and then for collaborative workshops incorporating both groups.

(4) In the second year, engagement with vertebrate pest professionals and wildlife managers also informed our planning for tool development. This was accomplished in part by a presentation by Stephanie Shwiff at the 31st annual Vertebrate Pest Conference in Monterey, California, March 11-14 2024.

(5) We attended the Maui Rangeland Health Workshop, on July 19, 2023. There were approximately 30 participants at the workshop hosted by Maui Cattlemen’s Association. We met with producers, State and Federal managers, and researchers regarding axis deer issues impacting ecosystem services, soil health, watershed function, aquifers, and coral reefs, as well as management for resilience to climate change. 

 

10 Producers reported gaining knowledge, attitude, skills and/or awareness as a result of the project
Key areas taught:
  • Pest management
Key changes:
  • Pest management

Information Products

    Any opinions, findings, conclusions, or recommendations expressed in this publication are those of the author(s) and should not be construed to represent any official USDA or U.S. Government determination or policy.